Associate Professor
Telephone: (01792) 295443
Room: Academic Office - 100
First Floor
Wallace Building
Singleton Campus

My research focusses on the impacts of stochastic environmental variability on population processes affecting ecological and evolutionary dynamics.

I combine analytical, simulation and statistical modelling approaches to investigate how different forms of environmental variation (spatial and/or temporal) drive changes in population size and behavioural decisions. Where possible, I also try to link these approaches to data from natural populations.

Areas of Expertise

  • Complex dynamics
  • environmental stochasticity
  • species–environment interactions
  • dispersal
  • species–species interactions.
  • coloured stochastic processes
  • synchrony
  • spatio-temporal dynamics
  • data analysis
  • quantitative approaches
  • Saltmarsh


  1. Mugabo, M., Gilljam, D., Petteway, L., Yuan, C., Fowler, M., Sait, S. Environmental degradation amplifies species' responses to temperature variation in a trophic interaction Journal of Animal Ecology 88 11
  2. Kolesidis, D., Boddy, L., Eastwood, D., Yuan, C., Fowler, M. Predicting fungal community dynamics driven by competition for space Fungal Ecology 41 13 22
  3. Astudillo-Sánchez, C., Fowler, M., Villanueva-Díaz, J., Endara-Agramont, A., Soria-Díaz, L. Recruitment and facilitation in Pinus hartwegii, a Mexican alpine treeline ecotone, with potential responses to climate warming Trees
  4. Alhadidi, S., Fowler, M., Griffin, J. Functional diversity of predators and parasitoids does not explain aphid biocontrol efficiency BioControl
  5. Yang, Q., Fowler, M., Jackson, A., Donohue, I. The predictability of ecological stability in a noisy world Nature Ecology & Evolution 3 2 251 259

See more...


  • BIO109 Core Skills for Biological Sciences

    This module is divided into three sections, scientific writing, data analysis and chemistry, which will equip students with the core skills needed throughout their degree program. The content of the module includes understanding the different types of data that can be measured and collected, the tools to formally present and analyse data and data analyses, and practical applications of spreadsheet software. There is a 'hands on' focus on dealing with data and developing basic mathematical and statistical analytical skills. Furthermore this module introduces first year undergraduates to the key skill of scientific writing, developing their ability to locate, understand, evaluate and communicate scientific information. Basic chemistry will be covered as a foundation to its importance to biological processes.

  • BIOM25B Science Skills and Research Methods

    This intensive lecture and practical based module covers science skills for students wishing to pursue postgraduate studies, including MSc, MRes, MPhil and PhD degrees. It will teach students how to make good use of library and internet resources (including Web of Science, Voyager and Blackboard), to design and analyse their experiments, and to make presentations of their data during conferences and symposia. It will provide PG students in the Department of Biosciences (and other Departments in the College of Science) with the research and analytical skills necessary to carry out their research projects. It will teach them how to formulate and test scientific hypotheses, and how to generate and analyse scientific results using a variety of research methods. Lecture topics include Reporting and Presentation skills, Numerical skills, Philosophy and Methodology of Science, and Biostatistics. The lectures are taught during the first part of the Semester. The module is examined through a combination of CA (50%) and Examination in the form of a MCQ test (50%). Basic reading: Whitlock, M. and Schluter, D. (2014) The Analysis of Biological Data (Roberts & Co.). Crawley, M.J. (2005) Statistics: An Introduction Using R (Wiley). Original research papers given in reading list


  • Assessment of Himalayan balsam (Impatiens glandulifera) removal from riparian zones and its impact on intra-river biodiversity and habitat quality: Implications for catchment level best management practices. (current)

  • Effects of Biodiversity Loss on Rockpool Communites (current)

    Other supervisor: Dr John Griffin
  • Exploring protein expression in antagonistic wood decay fungal interactions (current)

    Other supervisor: Prof Dan Eastwood
  • Resilience of ecosystem functions in vegetated costal systems: a trait-based perspective (awarded 2020)

    Other supervisor: Dr John Griffin
  • Trade-offs between multiple ecosystem services in UK and US salt marshes (awarded 2020)

    Other supervisor: Dr John Griffin
  • Using MaXENT to analyse the distribution of Himalayan balsam (Impatiens glandulifera) in Wales (awarded 2019)

    Other supervisor: Dr Daniel Forman
  • Rewiring evolutionary games: How does cooperation survive in a small world (awarded 2019)

  • The Forgotten Ecosystem Engineers: Community and Functional Ecology of Pioneer Wood Decay Fungi in the Canopy of Beech Trees (awarded 2019)

    Other supervisor: Prof Dan Eastwood
  • 'Inducible defences in two green algae as reponse to the predation by cladocerans' (awarded 2019)

    Other supervisor: Prof Kam Tang
  • Biodiversity of natural enemies and the control of agricultural pests (awarded 2019)

    Other supervisor: Dr John Griffin
  • Modelling direct biotic interactions with a focus on filamentous fungi (awarded 2018)

    Other supervisor: Dr Lloyd Bridge
    Other supervisor: Prof Chenggui Yuan
  • 'The Causes and Consequences of Variation in Different Dimensions of Biodiversity on Rocky Shores' (awarded 2018)

    Other supervisor: Dr John Griffin