Senior Lecturer
Telephone: (01792) 602199
Email: JavaScript is required to view this email address.

Areas of Expertise

  • Animal Colouration
  • Visual Ecology
  • Comparative Methods
  • Life History
  • Invasion Biology


  1. & Avoiding Attack: The Evolutionary Ecology of Crypsis, Aposematism, and Mimicry.. Oxford: Oxford University Press.
  2. et. al. The biology of color. Science 357(6350), eaan0221
  3. & Experimental evidence that primate trichromacy is well suited for detecting primate social colour signals. Proceedings of the Royal Society B: Biological Sciences 284(1856), 20162458
  4. & Interspecific visual signalling in animals and plants: a functional classification. Philosophical Transactions of the Royal Society B: Biological Sciences 372(1724), 20160344
  5. & Fast life history traits promote invasion success in amphibians and reptiles. Ecology Letters 20(2), 222-230.

See more...


  • BIO229 Tetrapod Evolution

    This module follows on from the introduction to vertebrates in the Level 4 Animal Diversity, Form and Function module, providing detail on form and function in vertebrates that spend all or part of their life cycle on land. Aspects of tetrapod behaviour, morphology and physiology will be considered in terms of adaptation and evolutionary constraint. Practicals will provide an introduction to the anatomy of birds and mammals by means of dissection, inference of the phylogenetic relationships between avian species, and an exploration of how beak morphology affects ecological niche in birds. Overall, students will gain an appreciation of the diversity of tetrapod types and an insight into the fundamental importance of metabolic rate in animals.

  • BIO253 Introduction to field zoology

    This residential field course comprises practical work employing techniques appropriate to sampling the zoological biodiversity of a range of terrestrial and freshwater habitats (coastland, woodland, grassland, freshwater systems). Students will learn techniques for the identification of species, practice recording accurate field notes, and gain experience in the analysis and presentation of zoological and ecological data. Furthermore students will be able to recognise and describe key groups of animals associated with a range of temperate habitats.

  • BIO342 Sensory Ecology

    Sensory Ecology aims to understand how organisms acquire, process and respond to information. It is a relatively new field within the biological sciences that draws on knowledge and techniques from a wide range of disciplines, beginning with the physics that defines the structure of information in the environment, through to the anatomy, neurophysiology, perceptual psychology and psychophysics that aims to understand how this information is sensed, processed and perceived. The subject also includes the behavioural ecology and evolutionary biology that seeks proximate and ultimate explanations for sensory systems. The module emphasises the interdisciplinary nature of studying sensory systems and aims to make links between different kinds of explanation.


  • Novel versatile semiochemical dispensers for improved pest monitoring and control (current)

    Student name:
    Other supervisor: Dr Joel Loveridge
    Other supervisor: Prof Tariq Butt

    Student name:
    Other supervisor: Dr Richard Unsworth
    Other supervisor: Dr Hanna Nuuttila
  • Untitled (current)

    Student name:
    Other supervisor: Prof Luca Borger
  • Welfare benefits of physical activity and microalgal feed supplementation in Zebrafish (Danio rerio) (current)

    Student name:
    Other supervisor: Prof Sofia Consuegra Del Olmo
    Other supervisor: Prof Carlos Garcia De Leaniz
  • Comparing the rate of weapon and ornament diversification. (current)

    Student name:
    Other supervisor: Dr Kevin Arbuckle
  • Distinct bacterial communities associated with Zostera marina roots may persist along ontogenetic gradients (current)

    Student name:
    Other supervisor: Dr Richard Unsworth
    Other supervisor: Dr Richard Unsworth
  • Camouflage during pursuit (awarded 2019)

    Student name:
    Other supervisor: Dr Emily Shepard

Career History

Start Date End Date Position Held Location
April 2016 Present Lecturer Swansea University

Administrative Responsibilities

  • MPhil/PhD Programme Coordinator - Biosciences

    2017 - Present