Senior Lecturer
Telephone: (01792) 602883
Email: JavaScript is required to view this email address.
Room: Office - 136
First Floor
Vivian Building
Singleton Campus

Areas of Expertise

  • Ocean acidification
  • Invertebrate biology
  • Fish biology
  • Invertebrate immunology
  • Larval biology
  • Cephalopoda
  • Biofouling


  1. & European sea bass, Dicentrarchus labrax, in a changing ocean. Biogeosciences 11(9), 2519-2530.
  2. & Enhanced Cellular Immunity in Shrimp (Litopenaeus vannamei) after ‘Vaccination’. PLoS ONE 6(6), e20960
  3. & Estimating the ecological, economic and social impacts of ocean acidification and warming on UK fisheries. Fish and Fisheries
  4. & European sea bass show behavioural resilience to near-future ocean acidification. Royal Society Open Science 3(11), 160656
  5. & Increased Disease Calls for a Cost-Benefits Review of Marine Reserves. PLoS ONE 7(12), e51615

See more...


  • BIO224 Ichthyology

    This module follows on from BIO105 Animal Diversity to discuss the evolution, ecology, structure and functional physiology of the paraphyletic group of animals referred to as fish. There will also be some detail on fisheries. Practicals will provide an introduction to the anatomy of fish by means of dissection and an appreciation of the diversity of fish types and their very different ecological roles. The aquarium visit will also introduce students to behavioural inventories and their role in determining animal welfare.

  • BIO237 Marine Invertebrates

    This module introduces students to the vast diversity of marine invertebrate and the fundamental roles they play in marine ecology. Students will receive 18 lectures and three laboratory practicals covering the general themes of: marine invertebrate taxonomy and developmental biology; form, function and behaviour; comparative physiology; reproductive strategies and biogeography; and ecological roles. Students will be examined on their understanding of the lecture material, recommended reading and practical techniques.

  • BIO251 Biosciences Year 2 field course alternative assessment

    This module is an alternative for students that are unable to attend the residential Biology, Zoology or Marine Biology field courses in Year 2. In order to qualify for this module, students need to have a satisfactory reason that has been authorised by the module coordinators before the field course is undertaken. Evidence of this will be required. Students will be supplied with data to analyse and directed to research and investigate relevant habitats that emulate those studied on the field course.

  • BIO260 Marine Biology Field Course

    This residential field course comprises practical work employing shore-based techniques to sample littoral and benthic marine habitats. Students will learn techniques for the identification of marine organisms and gain experience in the analysis and presentation of ecological data.

  • BIO327 Tropical marine ecology field course

    This field based module will provide students with an introduction to the ecology of tropical marine systems and teach students the key practical skills required by tropical marine biologists. Students will obtain training in how to design, implement and report scientifically robust marine research. The module will complement the level three marine field course and help develop key skills in field based marine biology. Students will learn skills in marine ecology and taxonomy, in-water marine sampling and surveys, and impact assessment. This module will be mostly practical based but will also include theory lectures, workshops and feedback sessions. It would be structured around seven days of directed practical activities and a three day small group based mini-project. The field course will utilise snorkeling and intertidal walking as the major means of sampling throughout directed practical¿s.

  • BIO346 Professional Skills in Marine Biology

    This field-based module will introduce students to the professional techniques utilised to monitor and study marine life in a variety of marine and coastal habitats and in relation to conservation management and biodiversity monitoring in the United Kingdom. The course places a strong emphasis on marine ecological census techniques. Students will learn key skills relevant to the marine ecology sector including protected and economically-important species (especially marine mammals, fish, shellfish, coastal birds), Phase 1 habitat surveys and water quality surveys. Students will also learn about the biotic and abiotic factors that define different UK habitats and relevant regulations that protect them. The module provides an introduction to the Environmental Impact Assessment (EIA) process and a range of impacts on the marine environment including energy generation and pollution. Participants in this module will work in groups acting as a marine environmental consultancy and the class will be responsible for producing key survey results for an Environmental Impact Assessment (EIA) and public engagement presentation for a proposed commercial development in Carmarthen Bay. Students will gain insider experience about professional techniques in marine (and freshwater) biology through a series of lectures delivered by marine environmental practitioners from environmental consultancies and regulatory organisations in the UK. The course includes a five-day residential field course in September/October which provides the students with the opportunity to practice the key technical skills in a real-world setting

  • BIO350 Biosciences Research Project

    This module is designed to develop the research and/or survey skills of undergraduate students in Biology, Marine Biology and Zoology. It covers literature reviewing; research planning and experimental/survey design; safety assessment; data collection techniques; data analysis and presentation; critical evaluation; discussion of results in the light of published work; final report production and presentation of results at a research symposium.


  • A novel approach to native oyster production and associated ecosystem services (current)

    Student name:
    Other supervisor: Dr Christopher Coates
  • Investigations into biotic growth and corrosion on tidal range marine hydropower turbines (awarded 2018)

    Student name:
    Other supervisor: Prof Ian Masters
  • '''''Spatial learning in a decapod crustacean''''' (awarded 2018)

    Student name:
    Other supervisor: Prof Mary Gagen
  • 'Macroalgae as a potential mitigation agent for ocean acidification' (awarded 2017)

    Student name:
    Other supervisor: Prof Kam Tang