Senior Lecturer
Telephone: (01792) 602883
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Areas of Expertise

  • Ocean acidification
  • Invertebrate biology
  • Fish biology
  • Invertebrate immunology
  • Larval biology
  • Cephalopoda
  • Biofouling


  1. & Estimating the ecological, economic and social impacts of ocean acidification and warming on UK fisheries. Fish and Fisheries
  2. & European sea bass show behavioural resilience to near-future ocean acidification. Royal Society Open Science 3(11), 160656
  3. Disease profiles of juvenile edible crabs (Cancer pagurus L.) differ at two geographically-close intertidal sites. Journal of Invertebrate Pathology 128, 1-5.
  4. & European sea bass, Dicentrarchus labrax, in a changing ocean. Biogeosciences 11(9), 2519-2530.
  5. Bacterial septicaemia in prerecruit edible crabs,Cancer pagurusL.. Journal of Fish Diseases 37(8), 729-737.

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  • BIO224 Ichthyology

    This module follows on from BIO105 Animal Diversity to discuss the evolution, ecology, structure and functional physiology of the paraphyletic group of animals referred to as fish. There will also be some detail on fisheries. Practicals will provide an introduction to the anatomy of fish by means of dissection and an appreciation of the diversity of fish types and their very different ecological roles.

  • BIO237 Marine Invertebrates

    This module introduces students to the vast diversity of marine invertebrate and the fundamental roles they play in marine ecology. Students will receive 18 lectures and three laboratory practicals covering the general themes of: marine invertebrate taxonomy and developmental biology; form, function and behaviour; comparative physiology; reproductive strategies and biogeography; and ecological roles. Students will be examined on their understanding of the lecture material, recommended reading and practical techniques.

  • BIO251 Biosciences Year 2 field course alternative assessment

    This module is an alternative for students that are unable to attend the residential Biology, Zoology or Marine Biology field courses in Year 2. In order to qualify for this module, students need to have a satisfactory reason that has been authorided by the module coordinators before the field course is undertaken. Evidence of this will be required. Students will be supplied with data to analyse and directed to research and investigate relevant habitats that emulate those studied on the field course.

  • BIO260 Marine Biology Field Course

    This residential field course comprises practical work employing shore-based techniques to sample littoral and benthic marine habitats. Students will learn techniques for the identification of marine organisms and gain experience in the analysis and presentation of ecological data.

  • BIO327 Tropical marine ecology field course

    This field based module will provide students with an introduction to the ecology of tropical marine systems and teach students the key practical skills required by tropical marine biologists. Students will obtain training in how to design, implement and report scientifically robust marine research. The module will complement the level three marine field course and help develop key skills in field based marine biology. Students will learn skills in marine ecology and taxonomy, in-water marine sampling and surveys, and impact assessment. This module will be mostly practical based but will also include theory lectures, workshops and feedback sessions. It would be structured around seven days of directed practical activities and a three day small group based mini-project. The field course will utilise snorkeling and intertidal walking as the major means of sampling throughout directed practical¿s.

  • BIOM51 Masters Literature Reviews (Biosciences)

    Students will learn how to write a scientific literature review to the level required by a peer-reviewed journal. Initially, they will produce a 4,000 word review (40% of final mark) which will require them to: 1) Devise a scientific question 2) Critically evaluate this question in the context of the available scientific literature 3) Revise the question as necessary 4) Perform a detailed review of the literature around this question 5) Synthesise the literature into an evidence-based narrative 6) Draw solid conclusions 7) Accurately and consistently reference literature These skills will be further developed by producing a second, 6,000 word review (60% of final mark) with a title chosen from a provided list. The mark and feedback on the first review will be designed to help the student improve their performance in this second piece of work. A workshop delivered by the students' tutor will help them develop their initial question and writing style. There are no examinations for this module.


  • A novel approach to native oyster production and associated ecosystem services (current)

    Student name:
    Other supervisor: Dr Christopher Coates
  • Investigations into biotic growth and corrosion on tidal range marine hydropower turbines (awarded 2018)

    Student name:
    Other supervisor: Prof Ian Masters
  • '''''Spatial learning in a decapod crustacean''''' (awarded 2018)

    Student name:
    Other supervisor: Prof Mary Gagen
  • 'Macroalgae as a potential mitigation agent for ocean acidification' (awarded 2017)

    Student name:
    Other supervisor: Prof Kam Tang