Areas of Expertise

  • Disease ecology - with particular reference to shellfish of commercial importance
  • Evolutionary immunology
  • Aquaculture


  1. & A novel bacterial infection of the edible crab, Cancer pagurus. Journal of Invertebrate Pathology 133, 83-86.
  2. & Effects of Experimental Infection of Juvenile Edible Crabs Cancer pagurus with the Parasitic Dinoflagellate Hematodinium sp.. Journal of Shellfish Research 34(2), 511-519.
  3. & Effects of population density and body size on disease ecology of the European lobster in a temperate marine conservation zone. ICES Journal of Marine Science: Journal du Conseil 72(suppl 1), i128-i138.
  4. & Disease profiles of juvenile edible crabs (Cancer pagurus L.) differ at two geographically-close intertidal sites. Journal of Invertebrate Pathology 128, 1-5.
  5. & How Does the Dinoflagellate Parasite Hematodinium Outsmart the Immune System of Its Crustacean Hosts?. PLOS Pathogens 11(5), e1004724

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  • BIO236 Cells and Immunity

    This module consists of the following lectures and practicals: Lecture Topics: 1. Advances in microscopy (2) 2. Cell, tissue and organ culture - bioengineering and stem cells (3) 3. Immunology (15) Practicals: 1. Live cells ¿ methods of observation and counting (1) 2. Human haematology (2) 3. Immunodiffusion (2)

  • BIO313 Diseases of aquatic animals

    This is a research-led module that explores recent advances in knowledge of diseases of aquatic organisms with particular reference to invertebrates (mainly crustaceans and molluscs) and fish of commercial significance

  • PM-140 Anatomy

    This module will provide knowledge of the structure of the human body, and how this adult anatomy develops. It will study human anatomy in a systems approach, focusing on the cardiovascular, gastrointestinal, musculoskeletal, urogenital, and nervous systems. Anatomy is a fundamental science and supports many areas of biology. As such the topics chosen for this module are those most useful to other areas of biological science, and are often clinically significant. This module will use practical classes to study human tissue and will develop the skills needed to dissect and study cadavers. Support materials and laboratory space for self directed learning will be available, including plastic anatomical models, bones and skeletons, and computer based anatomical models.


  • The pathogens and parasites of the common shore crab, Carcinus maenas (current)

    Student name:
    Other supervisor: Dr Christopher Coates
  • Immunobiology of Crepidula fornicata and its potential as a harbourer of infectious disease (current)

    Student name:
    Other supervisor: Dr Christopher Coates